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Dotarłem do chińskiej pracy: Min Zhang, Yang Zhong, Tianwen Cao, Yupeng Geng, Yuan Zhang, Ke Jin, Zhumei Rena, Rui Zhanga, Yaping Guoa, Enbo Maa: Phylogenetic relationship and morphological evolution in the subfamily Limenitidinae (Lepidoptera: Nymphalidae) Progress in Natural Science 18 (2008) 1357–1364, która w abstrakcie podaje:
zaś konkretnie odnośnie Aglais ioThe mitochondrial cytochrome oxidase subunit I (COI) gene and the nuclear elongation factor 1o (EF-1a) gene were sequenced from
29 species of Nymphalidae (Nymphalidae, Lepidoptera). Phylogenetic trees were constructed based on the sequences determined from
the 29 species and sequences of other 36 species deposited in GenBank using the neighbor-joining (NJ), maximum likelihood (ML) and
Bayesian methods with Libythea celtis (Libytheinae) as the outgroup. Our phylogenetic trees indicated four major clades. Clade A
includes three subfamilies: Apaturinae, Nymphalinae, and Limenitidinae, excluding the tribe Limenitidini; Clade B includes the subfam-
ilies Heliconiinae and the tribe Limenitidini; Clade C includes Satyrinae, Calinaginae, Charaxinae and Morphinae; and Clade D includes
subfamily Danainae. Our study suggested that the tribes Pseudergolini, Biblidini, Limenitidini and Cyrestidini should be considered as
subfamilies and confirmed the interspecific relationships within the subfamily Pseudergolinae, namely Amnosia + (Pseudergolis + (Stibo-
chiona + Dichorragia)). We then mapped three morphological characters (spot of anal angle, eyespots, and process from outer margin of
hind wing) onto the phylogenetic tree constructed by ML analysis using the combined sequence data. Based on this the evolutionary
patterns of these morphological characters were identified, they indicated that the three characters evolved repeatedly in the family
Nymphalidae.
P.S. jak widać to całe zamieszanie to wina ChińczykówThe phylogenetic positions of the species Aglais io and
Polygonia canace have long been controversial. Paul [7]
placed the species Aglais io and Polygonia canace in the
genus Vanessa. However, Chou [31,32], Li and Zhu [33]
and Bai et al. [34] suggested that the species Aglais io and
Polygonia canace should be included in the genus Inachis
and Kaniska, respectively. In this study, the species Aglais
io was found to be a sister to Aglais urticae with a strong
support (bootstrap value 94%), which agreed with Harvey’s
suggestion [6] that moving the species Aglais io into the
genus Aglais.